:root {
  --canvas: #f7f7f4;
  --canvas-soft: #fafaf7;
  --surface-card: #ffffff;
  --surface-strong: #e6e5e0;
  --hairline: #e6e5e0;
  --hairline-soft: #efeee8;
  --hairline-strong: #cfcdc4;
  --primary: #f54e00;
  --primary-active: #d04200;
  --ink: #26251e;
  --body: #5a5852;
  --muted: #807d72;
  --muted-soft: #a09c92;
  --on-primary: #ffffff;
  --success: #1f8a65;
  --error: #cf2d56;
  --t-thinking: #dfa88f;
  --t-grep: #9fc9a2;
  --t-read: #9fbbe0;
  --t-edit: #c0a8dd;
  --t-done: #c08532;
  --sans: "Inter", system-ui, "Helvetica Neue", Helvetica, Arial, sans-serif;
  --mono: "JetBrains Mono", ui-monospace, SFMono-Regular, Menlo, monospace;
  --maxw: 1200px;
}

* { box-sizing: border-box; }

html { scroll-behavior: smooth; }

body {
  margin: 0;
  background: var(--canvas);
  color: var(--body);
  font-family: var(--sans);
  font-size: 16px;
  line-height: 1.5;
  letter-spacing: -0.15px;
  -webkit-font-smoothing: antialiased;
}

.container { width: 100%; max-width: var(--maxw); margin: 0 auto; padding: 0 24px; }

a { color: var(--ink); text-decoration: none; }
a:hover { color: var(--primary); }

h1, h2, h3 { color: var(--ink); font-weight: 400; margin: 0 0 16px; }
h1 { font-size: 72px; line-height: 1.1; letter-spacing: -2.16px; }
h2 { font-size: 36px; line-height: 1.2; letter-spacing: -0.72px; }
h3 { font-size: 26px; line-height: 1.25; letter-spacing: -0.325px; }
p { margin: 0 0 16px; }

.skip-link {
  position: absolute; left: -9999px; top: 0; background: var(--ink);
  color: var(--canvas); padding: 10px 16px; border-radius: 8px; z-index: 200;
}
.skip-link:focus { left: 16px; top: 16px; }

.eyebrow {
  display: inline-block; font-size: 11px; font-weight: 600; line-height: 1.4;
  letter-spacing: 0.88px; text-transform: uppercase; color: var(--muted);
  background: var(--surface-strong); padding: 4px 10px; border-radius: 9999px;
  margin-bottom: 16px;
}

.top-nav {
  background: var(--canvas); border-bottom: 1px solid var(--hairline);
  position: sticky; top: 0; z-index: 100;
}
.nav-inner { display: flex; align-items: center; justify-content: space-between; height: 64px; }
.brand { display: inline-flex; align-items: center; gap: 8px; font-weight: 600; font-size: 18px; color: var(--ink); }
.brand:hover { color: var(--ink); }
.brand-mark { width: 14px; height: 14px; border-radius: 4px; background: var(--primary); display: inline-block; }
.nav-menu { display: flex; align-items: center; gap: 24px; }
.nav-link { font-size: 14px; font-weight: 500; color: var(--body); }
.nav-cta {
  background: var(--ink); color: var(--canvas); padding: 8px 16px;
  border-radius: 8px;
}
.nav-cta:hover { color: var(--canvas); background: #14130f; }
.nav-toggle { display: none; flex-direction: column; gap: 4px; background: none; border: 0; cursor: pointer; padding: 8px; }
.nav-toggle span { width: 22px; height: 2px; background: var(--ink); display: block; }

.btn {
  display: inline-flex; align-items: center; justify-content: center;
  font-family: var(--sans); font-size: 14px; font-weight: 500; cursor: pointer;
  border-radius: 8px; border: 1px solid transparent; padding: 10px 18px;
  height: 40px; transition: background .15s, color .15s;
}
.btn-primary { background: var(--primary); color: var(--on-primary); }
.btn-primary:hover { background: var(--primary-active); color: var(--on-primary); }
.btn-secondary { background: var(--surface-card); color: var(--ink); border-color: var(--hairline-strong); }
.btn-secondary:hover { color: var(--ink); border-color: var(--ink); }
.btn-download { background: var(--ink); color: var(--canvas); padding: 12px 20px; height: 44px; }
.btn-download:hover { background: #14130f; color: var(--canvas); }

.hero-band { padding: 80px 0; border-bottom: 1px solid var(--hairline); }
.hero-grid { display: grid; grid-template-columns: 1.05fr 0.95fr; gap: 48px; align-items: center; }
.hero-copy h1 { margin-bottom: 20px; }
.hero-sub { font-size: 18px; color: var(--body); max-width: 38ch; margin-bottom: 24px; }
.hero-actions { display: flex; gap: 12px; flex-wrap: wrap; align-items: center; }
.text-link { font-size: 14px; font-weight: 500; color: var(--ink); border-bottom: 1px solid var(--hairline-strong); padding-bottom: 2px; }
.hero-figure { margin: 0; border: 1px solid var(--hairline); border-radius: 12px; overflow: hidden; background: var(--surface-card); }
.hero-figure img { display: block; width: 100%; height: 420px; object-fit: cover; }
.hero-figure figcaption { font-size: 13px; color: var(--muted); padding: 12px 16px; border-top: 1px solid var(--hairline); }

.section { padding: 80px 0; border-bottom: 1px solid var(--hairline); }
.section-head { max-width: 60ch; margin-bottom: 32px; }
.section-head p { font-size: 16px; color: var(--body); }

.grid-3 { display: grid; grid-template-columns: repeat(3, 1fr); gap: 24px; }
.grid-2 { display: grid; grid-template-columns: repeat(2, 1fr); gap: 24px; }

.card {
  background: var(--surface-card); border: 1px solid var(--hairline);
  border-radius: 12px; padding: 24px;
}
.card h3 { font-size: 22px; letter-spacing: -0.11px; }
.card p { font-size: 16px; color: var(--body); margin-bottom: 12px; }
.card .text-link { font-size: 14px; }

.species-card { background: var(--surface-card); border: 1px solid var(--hairline); border-radius: 12px; overflow: hidden; }
.species-card img { display: block; width: 100%; height: 200px; object-fit: cover; }
.species-card .species-body { padding: 20px; }
.species-card h3 { font-size: 18px; font-weight: 600; letter-spacing: 0; margin-bottom: 8px; }
.species-card .latin { font-style: italic; color: var(--muted); font-size: 14px; }
.species-card p { font-size: 15px; color: var(--body); margin: 8px 0 0; }

.callout {
  background: var(--canvas-soft); border: 1px solid var(--hairline);
  border-left: 3px solid var(--primary); border-radius: 12px; padding: 24px;
}
.callout h3 { font-size: 18px; font-weight: 600; letter-spacing: 0; }

.fact-list { list-style: none; padding: 0; margin: 0; }
.fact-list li { display: flex; gap: 12px; padding: 12px 0; border-bottom: 1px solid var(--hairline-soft); font-size: 16px; }
.fact-list li:last-child { border-bottom: 0; }
.fact-list .fact-label { font-weight: 600; color: var(--ink); min-width: 150px; }

.spec-table { width: 100%; border-collapse: collapse; background: var(--surface-card); border: 1px solid var(--hairline); border-radius: 12px; overflow: hidden; }
.spec-table th, .spec-table td { text-align: left; padding: 12px 16px; border-bottom: 1px solid var(--hairline-soft); font-size: 15px; }
.spec-table th { background: var(--canvas-soft); color: var(--ink); font-weight: 600; }
.spec-table tr:last-child td { border-bottom: 0; }

.code-note {
  font-family: var(--mono); font-size: 13px; background: var(--surface-card);
  border: 1px solid var(--hairline); border-radius: 12px; padding: 20px; color: var(--ink);
  overflow-x: auto;
}

.contact-band { padding: 80px 0; background: var(--canvas-soft); border-bottom: 1px solid var(--hairline); }
.contact-grid { display: grid; grid-template-columns: 1fr 1fr; gap: 48px; }
.form-field { margin-bottom: 16px; }
.form-field label { display: block; font-size: 14px; font-weight: 600; color: var(--ink); margin-bottom: 6px; }
.form-field input {
  width: 100%; background: var(--surface-card); color: var(--ink);
  border: 1px solid var(--hairline-strong); border-radius: 8px; padding: 12px 16px;
  height: 44px; font-family: var(--sans); font-size: 16px;
}
.form-field input:focus { outline: 2px solid var(--primary); outline-offset: 1px; border-color: var(--primary); }
.form-status { font-size: 14px; margin-top: 8px; min-height: 20px; }
.form-status.success { color: var(--success); }
.form-status.error { color: var(--error); }
.contact-info .fact-label { min-width: 90px; }

article.prose { max-width: 760px; margin: 0 auto; }
article.prose .lead { font-size: 20px; color: var(--body); margin-bottom: 24px; letter-spacing: 0.08px; }
article.prose h2 { font-size: 36px; margin-top: 48px; }
article.prose h3 { font-size: 22px; letter-spacing: -0.11px; margin-top: 32px; }
article.prose p, article.prose li { font-size: 17px; color: var(--body); line-height: 1.65; }
article.prose figure { margin: 32px 0; border: 1px solid var(--hairline); border-radius: 12px; overflow: hidden; background: var(--surface-card); }
article.prose figure img { display: block; width: 100%; height: auto; }
article.prose figcaption { font-size: 13px; color: var(--muted); padding: 12px 16px; border-top: 1px solid var(--hairline); }
article.prose ul, article.prose ol { padding-left: 22px; }
article.prose li { margin-bottom: 8px; }
.article-meta { font-size: 13px; color: var(--muted); margin-bottom: 24px; }
.article-meta span { margin-right: 16px; }
blockquote.note { margin: 24px 0; padding: 16px 20px; background: var(--canvas-soft); border-left: 3px solid var(--t-done); border-radius: 8px; color: var(--ink); font-size: 16px; }

.related { margin-top: 64px; padding-top: 32px; border-top: 1px solid var(--hairline); }
.related h2 { font-size: 26px; letter-spacing: -0.325px; }

.breadcrumb { font-size: 13px; color: var(--muted); margin-bottom: 24px; }
.breadcrumb a { color: var(--muted); }
.breadcrumb a:hover { color: var(--primary); }

.page-hero { padding: 64px 0 32px; }
.page-hero h1 { font-size: 56px; letter-spacing: -1.4px; }

.footer { background: var(--canvas); padding: 64px 0 32px; }
.footer-grid { display: grid; grid-template-columns: 1.6fr 1fr 1fr 1fr 1fr; gap: 32px; }
.footer-head { font-size: 11px; font-weight: 600; letter-spacing: 0.88px; text-transform: uppercase; color: var(--muted); margin-bottom: 16px; }
.footer-link { display: block; font-size: 14px; color: var(--body); padding: 4px 0; }
.footer-note { font-size: 14px; color: var(--muted); margin-top: 12px; }
.footer-bottom { margin-top: 48px; padding-top: 24px; border-top: 1px solid var(--hairline); font-size: 13px; color: var(--muted); }
.footer-bottom .disclaimer { margin-top: 8px; color: var(--muted-soft); max-width: 80ch; }

.cookie-banner {
  position: fixed; left: 0; right: 0; bottom: 0; z-index: 150;
  background: var(--surface-card); border-top: 1px solid var(--hairline-strong);
}
.cookie-inner { display: flex; align-items: center; justify-content: space-between; gap: 24px; padding: 16px 24px; }
.cookie-inner p { margin: 0; font-size: 14px; color: var(--body); }
.cookie-actions { display: flex; gap: 12px; flex-shrink: 0; }

@media (max-width: 1024px) {
  h1 { font-size: 56px; letter-spacing: -1.4px; }
  .hero-grid { grid-template-columns: 1fr; gap: 32px; }
  .grid-3 { grid-template-columns: repeat(2, 1fr); }
  .footer-grid { grid-template-columns: repeat(3, 1fr); }
}

@media (max-width: 768px) {
  .nav-toggle { display: flex; }
  .nav-menu {
    display: none; position: absolute; top: 64px; left: 0; right: 0;
    flex-direction: column; align-items: flex-start; gap: 0;
    background: var(--canvas); border-bottom: 1px solid var(--hairline); padding: 8px 24px 16px;
  }
  .nav-menu.open { display: flex; }
  .nav-link { padding: 10px 0; width: 100%; }
  .nav-cta { width: 100%; }
}

@media (max-width: 640px) {
  h1 { font-size: 32px; letter-spacing: -0.6px; }
  h2 { font-size: 28px; letter-spacing: -0.4px; }
  .page-hero h1 { font-size: 34px; }
  .grid-3, .grid-2, .contact-grid { grid-template-columns: 1fr; }
  .footer-grid { grid-template-columns: 1fr 1fr; }
  .hero-figure img { height: 280px; }
  .cookie-inner { flex-direction: column; align-items: stretch; }
  .cookie-actions .btn { flex: 1; }
}